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Hereditary platelet function defects

Hereditary platelet function defects

 

Platelet physiology

§        Megakaryopoiesis is regulated by TPO (synthesised by liver)

§        TPO binds receptor, c-mpl on circulating platelets and BM megas and is then internalised and degraded

§        Total mass of cells thus determines free TPO levels and regulates megakaryopoiesis and platelet production

§        Mean life span of 10 days

§        Upon vessel wall damage

§        Adhesion

§        vWF facilitates initial adhesion via GP1b/IX/V complex

§        slows down platelets and allows binding via GPIIb/IIIa (via vWF) and GPIa/IIa (via collagen)

§        Reduced GP1b/X/V = Bernard Soulier

§        Reduced GPIIb/IIIa = Glanzmann’s thrombasthenia

§        Aggregation and secretion

§        Activated by a range of agonists

§        ADP, thrombin, thromboxane

§        Receptor stimulation results in G protein interactions that trigger intracellular metabolic pathways

§        Shape change via cytoskeletal changes

§        Release of alpha and dense granule contents

§        Stimulation of phospholipase A2, liberation of thromboxane A2 and activation of GP IIb/IIIa receptors

 

Suggestive history

§        ISTH

§        2 or more of the following without blood transfusion or

§        1 or more of the following with blood transfusion or

§        1 symptom recurring on 3 distinct occasions

§        Nose bleeding >10 minutes in absence of trauma

§        Cutaneous bleeding with no or minimal trauma

§        Prolonged bleeding from trivial wounds >15 minutes

§        Oral cavity, including tooth extraction, post tonsillectomy etc. bleeding requiring medical attention

§        Spontaneous GI bleeding

§        Menorrhagia or bleeding from other mucosal sites

§        Positive family history

§        2 or more distinct bleeding sites

§        A single bleed so severe as to require blood transfusion

§        Need to stop certain drugs before testing (aspirin, clopidogrel, NSAIDs, antihistamines, beta-blockers, penicillin / cephalosporins. Aminophylline and some foods e.g. garlic)

 

Investigation

FBC

§        +/- citrated sample

 

Blood film

§   Platelet size

 

Coagulation screen

§        VWD 2N

 

Platelet adhesion testing

PFA-100

§        Aspirates whole blood under constant vacuum through a capillary and a microscopic aperture within a membrane

§        Membrane is coated with collagen and either ADP or epinephrine

§        Platelet plug forms at the aperture and eventually occludes = closure time

§        Sensitive for Glanzman’s and Bernard Soulier

§        Limited sensitivity for storage pool disorders

§        Not sensitive to vascular-collagen disorders or fibrinogen levels/ fibrin generation

§        Neither bleeding time or PFA are a perfect screening test

§        Coll/ epi = primary screening test

§        Coll/ ADP = normal in aspirin

§        Affected by

§        Platelet count

§        Haematocrit

§        Diet

§        Aspirin

§        vWF levels – need to measure vWF levels if abnormal result

§        Abnormal tests should be repeated to exclude transient defect

 

[also Retention in glass column and Baumgartner’s technique]

 

Platelet aggregation

§        Assay based on measuring the decrease in light absorbance that occurs in platelet rich plasma when platelets aggregate.

§        Aggregometer is set with platelet poor plasma to demonstrate 100% light transmittance, and platelet rich plasma is used to set the baseline at 0%.

§        Different agonists are then added to separate test aliquots, and as platelets aggregate light transmittance increases and results are plotted on moving graph paper.

§        Agonists include ADP, adrenaline, collagen and ristocetin, (arachidonic acid U46619 (thromboxane receptor agonist), TRAP (thrombin receptor activating peptide), heparin (HIT))

§        Recommended that full dose response curves are obtained with each agonist

§        In thrombocytopenic samples (<120) it is best to adjust the control sample to the same platelet count or perform studies on washed platelets in which the count can be adjusted (neither technique is perfect)

§        Limited sensitivity for storage pool disorders (25% will have normal platelet aggregation)

§        ADP release

§        Low concentrations of ADP cause primary or reversible aggregation.

§        ADP initially binds its receptor and releases intracellular Ca which causes a shape change (reflected by a small initial change in absorbance)

§        Fibrinogen then adds to the cell to cell contact and reversible aggregation occurs.  At low concentrations of ADP the platelets may dissociate after this phase.

§        Higher concentrations of ADP cause an irreversible secondary wave aggregation which is associated with the release of dense and alpha-granules as a result of activation of the arachidonic acid pathway. 

§        The primary wave is obscured by the secondary wave and defects will be missed if only high concentrations are used.

§        Collagen

§        This results in a single wave of aggregation after a lag phase, which results from activation of the arachidonic acid pathway

§        Ristocetin

§        This reacts with vWF and the membrane receptor to induce platelets to clump together (agglutination) and does not activate any of the aggregation pathways.

§        Arachidonic acid

§        AA induces TXA2 generation and granule release even if there is a defect of agonist binding to the surface membrane or the phospholiase induced release of endogenous arachidonate.

§        Aggregation only impaired if further steps in the pathway are impaired such as inhibition of cyclooxygenae (aspirin effect).

§        Pitfalls

§        Centrifugation – red cell contamination may cause apparent incomplete aggregation

§        Time – for 30 minutes after PRP preparation, platelets are unresponsive to agonists

§        Platelet count – low counts may cause slow/ weak aggregation

§        pH - <7.7 inhibits aggregation, >8.0 enhances aggregation

§        mixing speed - <800 or >1200rpm slows aggregation

§        haematocrit

§        temperature

§        dirty cuvette/ air bubbles in cuvette

 

Platelet granular content and release

§        ATP/ADP content and release (bioluminescence)

§        Platelet factor V release (Quebec platelet syndrome)

§        Electron microscopy

§        Platelet granule defects

§        Changes in platelet ultrastructure eg MHY-9 defects (previously classified as May-Heggelin, Flechtner, Sebastian, Epstein syndromes)

 

Flow cytometry

§        Uses much smaller volume of bloods

§        Glycoprotein surface expression used to diagnose

§        Glanzman thombasthenia (GPIIb or IIIa/ CD41 or 61) and Bernard Soulier (GPIb/ CD42b)

§        Scott syndrome (annexin V binding)

§        Fibrinogen binding

§        Annexin binding (to phosphatidylcholine)

§        P-selectin surface expression

 

Genetic analysis

§        Wiskott Aldrich

§        Offered to some families with severe platelet function disorders to allow prenatal diagnosis

 

Repsonse patterns

 

ADP

Adrenaline

Collagen

Ristocetin

Platelet nucletoides

Diagnosis

 

N

N

N

Absent

Increased (large platelets)

Bernard Soulier

Nucleotides normal in vWD

Flow for GPIb

Abs

Absent

Absent

Primary aggregation

N

Glanzmann’s

Flow for IIbIIIa

Prim

Prim

Prim

N

N

Aspirin or COX deficiency

Can also use AA to distinguish

Prim/ absent = SPD

Red = asp

 

Prim (at high concentrations, full irreversible aggregation)

Prim

Prim

N

Reduced ADP

Storage pool defect  dense

Prim

Prim

Prim

Prim

N

Red cell contamination

Plasma never clears

Red

Red

Red

N

Reduced

Low platelets

Slow and weak aggregation

N

Abs

N

N

N

Absent/ abberant adrenaline receptor

Eg. MPD

Increased at low conc

Increased at low conc

Increased at low conc

Increased at low conc

N

Hyperaggregation

Check if there is spontaneous aggregation

 

 

 

 

 

 

 

 

 

Heredity

Platelet count

Platelet size

PFA 100

L-Epi

ADP

Col

Risto (RIPA)

AA

A23187

Adhesion

 

 

 

 

 

 

 

 

 

 

Pseudo vWD (increased affinitiy for GpIb)

AD

Low

N

Inc

N

N

N

INC

N

 

Bernard-soulier (absent GpIb)

AR

Low

Large

Inc

N

N

N

0

N

N

Aggregation

 

 

 

 

 

 

 

 

 

 

Glanzmann (abnormal IIbIIIa)

AR

N

N

Inc

0

0

0

N

0

0

Storage pool

 

 

 

 

 

 

 

 

 

 

d granule (HP / CHS)

AR

N

N

Inc or Norm

1

High dose, full irreversible aggregation

R

1

1/0

R

a-granule (GPS)

AR

Low

Large

Inc or Norm

N

N/0

0

N

N

 

Signalling

 

 

 

 

 

 

 

 

 

 

Cyclooxygenase deficiency (aspirin)

 

N

N

 

 

1

R

N

R/0

N/R

Thromboxane

AR

 

 

 

 

1/N

R

N

R/0

N

ADP

AR

 

 

 

 

Rapidly reversible

 

 

 

 

Phospholipid

 

 

 

 

 

 

 

 

 

 

Scott syndrome

AR

Normal 

Normal 

 

+

+

+

+

+

 

Ehlers-Danlos

 

N

N

N

N

N

N

N

N

N

 

 

 

Adhesion

Pseudo or platelet VWD

§   AD

§   Mutations in platelet GP1b resulting in increased sensitivity to ristocetin

§   Reduced HMW VWF multimers

§   Important to distinguish from type IIb VWD as treated with platelets rather than VWF concentrates

§   Distinguised from type IIb by performing RIPA with donor platelets

 

Bernard Soulier syndrome

§   Deficieny of platelet GP Ib/V/IX complex

§        Decreased binding of vWF – reduced adherence to endothelium esp. where high shear stress

§   Giant platelets

§   Thrombocytopenia – due to reduced platelet survival

§   Normal aggregation except reduced in ristocetin

§   Same pattern of results as vWD, but not corrected by addition of normal plasma

§   Increased platelet nucleotides – suggestive of large platelets (normal in vWD)

§   Flow for accurate platelet count and for GP1b

§   Carriers mostly asymptomatic

 

Aggregation

Glanzmann’s thombasthenia

§        Fail to form aggregates because of reduced GPIIb/IIIa (CD41/ CD61)

§        Type 1 and type 2

§        Type 1: <10% GP IIb/IIIa complexes and absent platelet fibrinogen

§        Type 2: up to 30% GP IIb/IIIa with reduced platelet fibrinogen

§        Prolonged PFA100

§        Abnormal aggregation in all except ristocetin

§        In ristocetin, may have a primary response only

§        Confirm diagnosis by flow for GP IIb/IIIa (absent CD41 and 61 on flow)

§        Treat with platelet transfusions

§        Allo-immunisation and Abs to GP IIb/IIIa may develop

§        rVIIa can be used to arrest bleeding

§        Acquired form due to auto-Abs to GPIIb/IIIa (rare)

 

Storage pool defects

§        Either deficiency of granules or failure of normal secretion

Dense granule disorders (d SPD)

Dense granules containATP, ADP, calcium pyrophosphate and serotonin

§        Hermansky-Pudlak syndrome

§        AR

§        Oculocutaneous albinism

§        Total absence of dense bodies

§        Commonest genetic disorder in Puerto Rico

§        Can treat with DDAVP

§        Chediak-Higashi syndrome

§        AR

§        Oculocutaneous albinism, infections, LPD and neutrophil peroxidase positive inclusions

§        Classical finding is very large peroxidase-positive cytoplasmic granules in neutrophils

§        Death in first decade

§        Reduced or irregular dense bodies

§        Idiopathic deficiency/ δ–storage pool disease

 

α-granule disorders (CD62P = P-selectin)

§        Alpha-granules contain PDGF and PF4

§        Grey platelet syndrome

§        Large and agranular, thrombocytopenia

§        Contents of alpha granules are absent eg. PF4, PDGF, VWF, fibrinogen

§        Platelet nucleotides are normal

§        EM demonstrates reduced or absent granules

§        Aggregation – reduced response to collagen and ADP

§        Associated with myelofibrosis

§        P selectin retained and flow will be positive for this if platelets activated first

§        Platelets appear grey on blood film

§        Treate with DDAVP +/- platelets

§        Paris-Trousseasu or Jacobson syndrome

§        AD, thrombocytopenia

§        Giant abnormal alpha granules – can’t release their contents normally

§        Associated mental retardation, cardiac abnormalities, cranio-facial abnormalities

§        Quebec platelet syndrome or factor V quebec

§        AD – very rare

§        Low factor V within platelet α-granules, but normal plasma levels

§        Defective procoagulant activity due to failure in assembly of prothrombinase complex

§        Urokinase plasminogen is released in large quantities upon plt activation, therefore

§        Unresponsive to platelet transfusion, treat with anti-fibrinolytics

§        No characteristic aggregometry pattern

§        Arthrogryposis-renal dysfunction-cholestasis

 

Combined alpha and dense granule disorders

§   Rare - gene defect currently unknown

 

Platelet receptors and signal transduction pathways

§   Normal platelet count and morphology

§   Decreased primary aggregation to one or more agonists, accompanied by the absence of secondary aggregation to some or all

§   Nucleotide content and release will help to distinguish between SPDs

Thrombaxane A2 receptor defects

§        Impaired aggregation to arachidonic acid with preserved ristocetin response

§        Defects in COX or thromboxane synthase produce a similar pattern, but have preserved aggregation to prostaglandin or synthetic thromboxane U46619

ADP receptor defects

§        Aggregation to ADP is reduced and rapidly reversed

§        Due to mutations in ADP receptor P2Y12 which is needed for large aggregates to form (P2Y1 receptor is involved initially)

§        Clopidogrel produces the same phenotype (P2Y12 is the clopidogrel target)

Collagen receptor defects

§        Defects in GPVI  or GPIa/IIa

§        10 cases described

Adrenoreceptor defects

§        Absence of aggregation to adrenaline

Signalling

§        Not well characterised

§        Post receptor signalling pathways

§        Defects in G-protein subunits and phospholipase C enzymes

§        Many of these enzymes are expressed in multiple tissues but seem to be relatively deficient in the platelet

 

Phospholipid surface

Scott syndrome

§        AR

§        Primary haemostatic function is intact but there is failure of scramblase activity once platelet activated

§        Allows phosphatidylserine and phosphatidylenthanolamine to move from the inner to the outer membrane

§        Platelets have a reduced ability to promote factor X and prothrombin activation

§        Due to decreased surface exposure of phosphatidylserine plus a reduced shedding of microvesicles

§        Aggregation is normal

§        Flow to demonstrate absent PS on surface of activated platelets

§        Prothrombin consumption index test

 

Hereditary macrothrombocytopenias – MYH9 mutations

§        AD

§        MYH9 mutations are the most common (encodes non-muscle myosin II-A heavy chain – involved in cytoskeleton in megas, platelets and other tissues)

§        Previously classified as:

§        May-Heggelin anomaly

§        Fechtner syndrome

§        Sebastian syndrome

§        Epstein syndrome

§        Thrombocytopenia (20-130) with very large platelets and varying platelet dysfunction (usually mild)

§        Often associated with Granulocyte inclusions (Dohle-like bodies)

§        Phenotype very variable  may be associated with other congenital abnormalities including:

§        Glomerulonephritis (urinalysis should be part of the workup)

§        Sensorineural deafness

§        Cataracts

§        Electron microscopy

§        Definitive diagnosis requires demonstration of a genetic defect in MYH9

§        Management – variable as phenotype varies greatly – depends on personal and family history

§        As for other mild platelet disorders

 

Congenital amegakaryocytic thrombocytopenia

§        AR

§        Diagnosis

§        Severe congenital thrombocytopenia with virtually absent amegakaryocytes in marrow is highly suggestive

§        Mutations in MPL gene required for definitive confirmation (mutations in MPL result in abnormal function or expression of the thrombopoietin receptor)

§        Management

§        Platelet transfusion

§        Stem cell transplant – progresses to aplastic anaemia

Amegakaryocytic thrombocytopenia with radioulnar syntosis

§   AD – only described in a small number of families

§   Due to mutations in HOX A11

§   Management – platelet support + HSCT

 

Thrombocytopenia with absent radii

§        AR (or AD)

§        Molecular basis unknown

§        Severe thrombocytopenia

§        Bilateral absent radii

§        Cow’s milk intolerance, renal and cardiac abnormalities, facial capillary haemangiomas

§        Platelet count improves with age

§        Some develop AML in later life

§        Diagnosis

§        Distinctive clinical features

§        Reduced megakaryocytes in the bone marrow with abnormal maturation

§        Management

§        Platelet transfuions in infancy – usually not needed beyond this

§         

 

 

Wiskott-Aldrich Syndrome

§        X-linked recessive disease, WAS gene

§        Microthrombocytopenia, eczema and immunodeficiency (T and B cell)

§        AI disorders esp haemolytic anaemia and vasculitis

§        Malignancy (usually lymphoreticular)

§        Treat with HLA matched, irradiated and CMV neg platelets

§        Definitive treatment = BMT

 

Management

§        Registered with haemophilia centre

§        Green card etc

§        Life style advice / avoid NSAIDs etc

§        Tranexamic acid

§        Including gauze soaked in

§        DDAVP

§        Usually works for storage pool defects

§        Not usually for Glanzmann’s, Scott, occasional severe form of BSS

§        Platelets

§        Allo-immunisation or Abs to their missing GP

§        HLA matched

§        rFVIIa

§        licensed for GT where platelet refractoriness has been demonstrated

§        Babies of women with severe bleeding disorders eg. BSS, GT may be at risk of alloimmune thrombocytopenia if mum has platelet antibodies

§        Monitored for HLA and anti GPIIb/IIIa and GPIb antibodies

§        Plasma exchange has been used during pregnancy to reduce antibody titre

§        Transplantation may be appropriate

§        WAS

 

Acquired platelet dysfunction

§        Drugs

§        Aspirin – irreversible acetylation of platelet COX-1 and resulting inhibition of thromboxane A2 synthesis

§        Effect lasts as long as the platelet

§        NSAIDs – reversible inhibition of COX-1 (normal platelet function restored rapidly

§        Clopidogrel – bind irreversibly to the platelet purinergic receptor P2Y12 and inhibit platelet responses to ADP

§        American College of cardiology recommends cessation of aspirin 7-10days and clopidogrel 5 days prior to surgery

§        Aciximab – GPIIb/IIIa antagonist, inhibit platelet aggregation.

§        Bleeding in about 10%, but ICH and death are rare

§        Abciximab disassociates more slowly than others (eg. Tirofibran)

§        β-lactam antibiotics – most severe with low albumin

§        more unbound drug to interact with platelet surface

§        contribute to bleeding when there is a co-existing haemostatic defect

§        Uraemia

§        Increased NO impairs platelet-vessel wall interactions

§        Accumulation of guandinosuccinic acid which is the donor for NO

§        Improved with dialysis because of removal of GSA

§        Anaemia, because reduced axial flow means less platelet contact with vessel wall

§        Cirrhosis

§        Myeloma

§        PPs get adsorbed to platelet surface and cause dysfunction

§        Can also get acquired VWDS

§        Plasmapheresis may be helpful

§        Myeloproliferative disorders

§        Aquired VWD due to loss of HMWM

§        Increased platelet numbers deplete VWF

§        DDAVP doesn’t correct, cytoreduction is most appropriate treatment

§        Also associated with aortic stenosis

§        Cardiopulmonary by-pass

§        Blood bank platelets


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